Dr. Danilo Harms

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2026

• 2026/03

  Chitimia-Dobler, L., Mey, C., Harms, D., Hammel, J., Dunlop, J.A., Kotthoff, U., Mans, B.J.

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  Redescription of Alloceraea cretacea (Acari: Ixodida) with an additional nymphal fossil added to this species: Novel insights into the evolution of the Haematobothrion

  Parasitology

  https://doi.org/

• 2026/01

  JEONG, K., CHOI, Y., HARMS, D., YOO, J., KIM, S.

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  Two new species of Parobisium (Pseudoscorpiones: Neobisiidae) Chamberlin, 1930 from South Korea, with a preliminary phylogeny of the genus

  Zootaxa, 2, 5741

  https://doi.org/

2025

• 2025/12

  Lin, H., Harms, D., Cheng, R.

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  A temperate group living in the subtropics: evolutionary history and integrative taxonomy of the pseudoscorpion genus Allochthonius Chamberlin, 1929 (Pseudoscorpiones: Pseudotyrannochthoniidae) in Taiwan

  Organisms, Diversity & Evolution

  https://doi.org/

• 2025/11

  MILDE, M., FERNANDES MARTINS, T., HARMS, D., HAMMEL, J.U., DUNLOP, J.A., KOTTHOFF, U., CHITIMIA-DOBLER, L.

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  A new species of the Recent genus Amblyomma (Parasitiformes: Ixodidae) discovered in Dominican amber

  Palaeoentomology, 6, 8

  https://doi.org/

• 2025/10

  Michalik, P., Vink, C.J., Ramírez, M.J., Dederichs, T.M., Sirvid, P.J., Pawson, S.M., Harms, D.

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  Alone no more—Integrative taxonomy of odd‐clawed spiders challenges the monotypy of Pianoa and Gradungula ( )

  Systematic Entomology

  https://doi.org/

• 2025/09

  Harms, D., McRae, J., Curran, M., Harvey, M.S.

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  Identifying refugia for invertebrate conservation in biodiversity hotspots: examples from a new genus of dragon pseudoscorpions (Pseudotyrannochthoniidae: Karrichthonius)

  Invertebrate Systematics, 9, 39

  https://doi.org/

• 2025/08

  Hagen, M., Kotthoff, U., Harms, D., Loria, S.

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  New species of Lechytia Balzan, 1892 (Arachnida: Pseudoscorpiones) from Burmese amber highlights 99 million years of morphological stasis

  Cretaceous Research, 172

  https://doi.org/

• 2025/07

  Jeong, K., Harms, D., Yoo, J., Kim, S.

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  Four new species of dragon pseudoscorpions (Pseudoscorpiones: Pseudotyrannochthoniidae: Spelaeochthonius) from caves in South Korea revealed by integrative taxonomy

  PloS one, 7, 20

  https://doi.org/

• 2025/05

  Krämer, J., Lüddecke, T., Harms, D., Hurka, S., Predel, R.

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  Toxins Under a Rock: Proteo-transcriptomic Analysis Reveals the Venom Composition of the Pseudoscorpion Ammogarypus lawrencei (Pseudoscorpiones: Garypidae)

  Toxicon

  https://doi.org/

• 2025/03

  Harms, D., Melcher, A.C., Krehenwinkel, H., Birkhofer, K.

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  Estimating Required Sample Sizes for Gut Content Metabarcoding Studies of Dietary Diversity in Spiders

  Journal of Applied Entomology

  https://doi.org/

• 2025/01

  Harms, D., Kinga, H., Damptey, F.G., Jocqué, R., Henrard, A., Birkhofer, K.

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  Active restoration of post-mining forest benefits the activity density, but not the diversity of spider communities across the seasons in Ghana

  African Invertebrates, 66

  https://doi.org/

• 2025/01

  Loria, S.F., Frank, S., Dupérré, N., Smith, H.M., Jones, B., Buzatto, B.A., Harms, D.

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  The world’s most venomous spider is a species complex: systematics of the Sydney funnel-web spider (Atracidae: Atrax robustus)

  BMC Ecology and Evolution, 1, 25

  https://doi.org/

2024

• 2024/12

  Melcher, A.C., Weber, S., Birkhofer, K., Harms, D., Krehenwinkel, H.

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  To pool or not to pool: Pooled metabarcoding does not affect estimates of prey diversity in spider gut content analysis

  Ecological Entomology, 6, 49

  https://doi.org/

• 2024/11

  Neethling, J.A., Harms, D.

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  Afrogarypus foordi sp. nov. – a new pseudoscorpion species (Pseudoscorpiones, Geogarypidae) from South Africa

  African Invertebrates, 2, 65

  https://doi.org/

• 2024/08

  Melcher, A.C., Weber, S., Calaor, J., Kennedy, S., Gajski, D., Hörrmann, T., Schmidt, A., Melzl, L., Meiss, F., Hans, J., Mahla, L., Stothut, M., Harms, D., Birkhofer, K., Economo, E., Rogers, H., Gillespie, R., Roderick, G., Krehenwinkel, H.

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  Spiders as natural DNA samplers can recover arthropod community diversity and biotic interactions across space and time

  https://doi.org/

• 2024/06

  Röschmann, L.M., Harvey, M.S., Hou, Y., Harms, D., Kotthoff, U., Hammel, J.U., Ren, D., Loria, S.F.

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  First fossil species of family Hyidae (Arachnida: Pseudoscorpiones) confirms 99 million years of ecological stasis in a Gondwanan lineage

  PeerJ, 12

  https://doi.org/

• 2024/06

  Harms, D., Harvey, M.S., Roberts, J.D., Loria, S.F.

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  Tectonically driven climate change and the spread of temperate biomes: Insights from dragon pseudoscorpions (Pseudotyrannochthoniidae), a globally distributed arachnid lineage

  Journal of Biogeography, 6, 51

  https://doi.org/

• 2024/05

  Harms, D., Yoo, J.S.

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  The pseudoscorpion genus Nipponogarypus (Pseudoscorpiones, Olpiidae) found in seashore habitats in Japan and Korea

  Zoosystematics and Evolution, 3, 100

  https://doi.org/

• 2024/04

  Jeong, K., Harms, D., Yoo, J.

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  New species of the pseudoscorpion genus Pararoncus (Pseudoscorpiones, Syarinidae) from Korea

  Evolutionary Systematics, 1, 8

  https://doi.org/

• 2024/02

  Michalik, P., Smith, H.M., Milledge, G., Harms, D.

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  Another ghost of Gondwana—Progradungula barringtonensis Michalik & Smith, sp. nov., a new species of the relict spider genus Progradungula (Araneae: Gradungulidae) from a temperate rainforest in eastern Australia

  Austral Entomology, 1, 63

  https://doi.org/

• 2024/01

  Jeong, K., Harms, D., Johnson, J.

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  A new species of Ditha (Pseudoscorpiones, Chthoniidae, Tridenchthoniinae) from the Western Ghats of India, with an identification key for the genus

  Zoosystematics and Evolution, 1, 100

  https://doi.org/